Order Nassellaria

Family Theoperidae

Artostrobus annulatus (Bailey) (Figure 15.125). Cephalis cup-shaped, poreless, with a thin apical horn. Thorax cylindrical, with circular (proximally) to subrectangular (distally) pores in transverse rows, increasing in size toward the base. Shell height: up to 160 µm. Ref: Riedel (1958), Petrushevskaya (1967).

 

Clathrocyclas cassiopeiae Haeckel (Figure 15.126). Small cephalis with a stout apical horn and many smaller spines. Thorax large, campanulate, with irregular pores. Abdomen short, truncated, with ragged, spiny termination. Shell height: 100-200 µm. Ref: Haeckel (1887).

 

Cornutella profunda Ehrenberg (Figure 15.127). Narrow, bilocular conical shell with very small subspherical poreless cephalis with or without apical horn. Pores on thorax circular, increasing in size toward the base. Sell height: 120-230 µm. Ref: Riedel (1958), Nigrini (1967).

 

Corocalyptra cervus (Ehrenberg) (Figure 15.128). Cephalis with a large apical horn, often forked distally. Thorax large, campanulate, with regular, polygonal pores in transversal rows increasing in size distally. Abdomen restricted to a narrow brim with several rows of small pores. Shell height: 100-200 µm. Ref: Benson (1966).

 

Corocalyptra columba (Haeckel) (Figure 15.132). Subspherical cephalis partly submerged into cupola-shaped thorax, provided with a large apical horn. Thorax with three small wings. Abdomen cylindrical. Shell height: ca. 100-120 µm. Ref: Haeckel (1887), as Pterocorys columba.

 

Corocalyptra kruegeri Popofsky (Figure 15.136). Generally similar to C. columba; cephalis larger, thorax and abdomen cylindrical. Shell height: ca. 80 µm. Ref: Popofsky (1913).

 

Cycladophora davisiana (Ehrenberg) (Figure 15.131) [=Theocalyptra davisiana, ?Artostrobus jorgenseni]. Shell conical to campanulate. Cephalis subspherical, sparsely perforated, usually with two spines. Thorax conical to cylindrical, with circular (proximally) to quadrate (distally) pores increasing in size toward the base. Abdomen, when present, flared out, wider than thorax, with quadrate pores. Petrushevskaya (1967) described three subspecies of this form, of which C. davisiana (Ehrenberg) davisiana Petrushevskaya (Figure 15.131b) and C. davisiana (Ehr.) cornutoides (Petrushevskaya) (Figure 15.131c) are often used in current literature (the last one is probably synonymous with Artostrobus jorgenseni). Shell height: ca. 100 µm. Ref: Riedel (1958), Petrushevskaya (1967), Bjørklund and Ciesielski (1994).

 

Cyrtopera laguncula Haeckel (Figure 3O; 15.151) [=Cyrtolagena laguncula, Stichopera pectinata]. Very typical multisegmented shell with conical outline and clearly marked constrictions; last segment may be open or closed. Shell height: 150-250 µm. Ref: Petrushevskaya (1971a).

 

Dictyocephalus papillosus (Ehrenberg) (Figure 15.135) [=Carpocanarium papillosum]. Two-segmented, thick-walled shell. Cephalis spherical. Thorax oval, with circular, usually framed pores, with three short wings, ending in a narrowed, poreless peristome. Shell height: 70-90 µm. Ref: Petrushevskaya (1967).

 

Dictyophimus gracilipes Bailey (Figure 15.142) [=Pseudodictyophimus gracilipes, Dictyophimus clevei]. Cephalis partly submerged into thorax, bears a large apical horn. Thorax pyramidal or conical, mouth open or closed, with three conspicuous legs (dorsal and lateral spines). Shell height (without feet): 55-90 µm. Ref: Petrushevskaya (1971a).

 

Dictyophimus hirundo (Haeckel) (Figure 15.140) [=Pterocorys hirundo]. Cephalis, globular, sometimes spiny, with a stout, three-bladed apical horn. Thorax truncate-conical to campanulate, spiny, with large circular pores and three ribs which extend into massive, divergent feet. Highly variable species (species group?). Shell height (without feet): 50-100 µm. Riedel (1958), Nigrini and Moore (1979).

 

Dictyophimus infabricatus Nigrini (Figure 3Q; 15.143). Similar to D. hirundo, but shell is thinner and pores are larger; cephalis usually wih two horns (apical and vertical spines), feet are smaller. Shell height (without feet): 90-200 µm. Ref: Nigrini and Moore (1979).

 

Eucyrtidium acuminatum (Ehrenberg) (Figure 15.137) [=?Eucyrtidium hexagonatum]. Small subspherical cephalis with apical horn. Thorax small, inflated, thick-walled. Abdomen and four-five post-abdominal segments thin-walled, with pores arranged in longitudinal rows. Shell height: 120-200 µm. Eucyrtidium hexagonatum Haeckel is closely related to this species; according to Nigrini (1967) it can be distinguished from E. acuminatum by the sharp change in contour at the lumbar stricture. Ref: Nigrini and Moore (1979).

 

Eucyrtidium anomalum (Haeckel) (Figure 15.138). Cephalis spherical, partly submerged into the large, assymetric, conical, thin-walled thorax. Abdomen and two-three postabdominal segments thin-walled, with pores in longitudinal rows. Maximum shell width: 80-100 m. Ref: Petrushevskaya (1971a).

 

Eucyrtidium hexastichum (Haeckel) (Figure 15.139). Cylindrical or cnical shell with up to 9-10 segments, thin-walled, with pores arranged in transversal rows. Maximum shell width: 65-80 µm. Ref: Petrushevskaya (1971a).

 

Lipmanella bombus (Haeckel) (Figure 15.133) [=Dictyoceras cf. pyramidale]. Very characteristic thorax where the dorsal and main lateral spines, which project down and sideways, are surmounted by latticed keels; these keels define the three slightly concave sides of the pyramidal thorax. Maximum shell width: 80-150 µm. Ref: Benson (1966), Petrushevskaya (1971a).

 

Lipmanella dictyoceras (Haeckel) (Figure 3R, 15.134) [=Lipmanella virchowii, Dictyoceras virchowii, Dictyoceras neglectum]. Cephalis large, hemispherical, wih a conspicuous apical horn. Thorax thin-walled, conical-inflated, with three wings (dorsal and main lateral spines). Abdomen absent or rudimentary. Shell height: 90-130 µm. Ref: Petrushevskaya (1971a).

 

Litharachnium tentorium Haeckel (Figure 15.148). Two-segmented, very characteristic shell. Cephalis very small, spherical, hyaline. Thorax conical proximally, flaring rapidly outward distally and, in complete specimens, ending in a gently curved brim. Diameter of fully-grown shells: up to 1 mm. Ref: Benson (1966), Petrushevskaya (1971a).

 

Lithopera bacca Ehrenberg (Figure 15.129). Spherical, often rough cephalis with an eccentrically located apical horn partly submerged into an oval thorax with closed mouth. Pores on thorax regularly arranged. Shell height: 120-140 µm. Ref: Benson (1966).

 

Lithostrobus hexagonalis Haeckel (Figure 15.141). Multisegmented, subconical shell with 5-9 joints with well marked strictures and angular shoulders. Pores very regular, subpolygonal, in transverse rows. Shell height: 120-300 µm. Ref: Benson (1966).

 

Perypiramis circumtexta Haeckel (Figure 15.130) [=?Plectopyramis dodecomma]. Cephalis very small, ovate, hyaline. Thorax large, conical, with ca. 10 well defined longitudinal rows of subquadrate pores rapidly increasing in size toward the base. Pores are usually not aligned transversely. The form described as Plectopyramis dodecomma Haeckel differs from P. circumtexta in that pores are aligned transversely as well as longitudally. Bathropyramis woodringi is also similar to the above, but has thicker longitudinal bars and aligned horizontal (transverse) bars. Shell height: 100-300 µm. Ref: Riedel (1958), Nigrini and Moore (1979).

 

Pterocanium elegans (Haeckel) (Figure 15.147). Cephalis relatively small, with two large horns (apical and vertical spines). Thorax a large, thin-walled, three-sided pyramid with small, very regular subcircular pores and thin bars; edges of pyramid continue as three strong three-bladed legs. Abdomen cylindrical, may be rudimentary. Shell height (without horns): 180-260 µm. Ref: Benson (1966), as Pterocanium cf. elegans.

 

Pterocanium praetextum (Ehrenberg) group? (Figure 15.146). Similar to P. trilobum, from which it sometimes is difficult to separate; differs by having a thorax with more pronounced, angular shoulders, and often a better developed abdomen. Shell height (without horn and feet): 100-120 µm. Ref: Petrushevskaya (1971a).

 

Pterocanium trilobum (Haeckel) (Figure 3H; 15.145). Cephalis relatively small, with a stout apical horn. Thorax an inflated tetrahedron with regularly arranged circular pores, with three ribs extending into stout, three-bladed, slightly curved feet. Abdomen, when present, rudimentary. Shell height (without horn and feet): 100-120 µm. Ref: Nigrini and Moore (1979).

 

Pterocyrtidium dogieli Petrushevskaya (Figure 15.144) [=Sethoconus dogieli]. Cephalis hemispherical, almost poreless, thick-walled. Thorax cylindrical, thorny, with very large, irregular, subcircular pores. Shell height: ca. 140 µm. Ref: Petrushevskaya (1971a).

 

Sethoconus anthocyrtis Haeckel (Figure 15.150) [=Conarachnium polyacanthum, Lophocorys polyacantha]. Cephalis spherical, spiny. Thorax clearly differentiated from cephalis, very large, conical, spiny, with large, regular, subpolygonal pores approximately in longitudinal rows; termination ragged. Shell height: 200-300 µm. Ref: Haeckel (1887).

 

Theopilium tricostatum (Haeckel) (Figure 15.149) [=?Theocalyptra gegenbauri]. Chinese peasant hat-like shell with a small cephalis. Thorax widely open, with small, regularly arranged pores, circular proximally and becoming larger and more polygonal distally, with three conspicuous symmetrical ribs (dorsal and main lateral spines). Abdominal brim flat, with 5-8 rows of very regular, quadrangular pores. Shell diameter: 130-300 µm. Ref: Haeckel (1887), Benson (1966).